Footnotes

• Correction notice This article has been corrected since it was published Online First. The provenance and peer review statement has been amended.

• Contributors GDMC: carried out the histological analyses, reviewed and revised the manuscript, and approved the final manuscript as submitted. AIT: carried out the initial analyses, reviewed and revised the manuscript, and approved the final manuscript as submitted. JHM: conceptualised and designed the study, drafted the initial manuscript and approved the final manuscript as submitted.

• Competing interests None declared.

• Provenance and peer review Not commissioned; externally peer reviewed.